DIVULGACIÓN Y CIENCIA POPULAR / DIFFUSION AND POPULAR SCIENCE
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Total: 185 (1994: 1; 1995: 3; 1996: 2; 1997: 1; 1998: 2; 1999: 2; 2000: 1; 2004: 148; 2005: 11; 2006: 1; 2008: 10; 2009: 3)
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Copyright © 1999 - 2009 D. F. Cisneros-Heredia Sitio creado en / site created on: Noviembre / November 1999. Ultima modificación / Last change: 8 Noviembre / November 2009.
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Publicaciones completas disponibles gratis en formato pdf o html. Aviso: Aunque algunos articulos colocados en este sitio web son publicados bajo formato de acceso publico (open access), otros pueden ser descargados solo para uso personal, pero no copiados, reproducidos o vendidos a terceros.
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Full-text publications available free on PDF format or html. Note: Although some files made available in this web site are published under open access concepts, others are for personal use only, and can not be copied, reproduced or sold to third parties.
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Diego Francisco CISNEROS-HEREDIA
BIODIVERSIDAD & CONSERVACION / BIODIVERSITY & CONSERVATION
Cisneros-Heredia, D.F. (2009) Corales de Galápagos: La USFQ trabaja por la conservación del laboratorio
viviente de la evolución. Aula Magna (Publicación Quincenal de la Universidad San Francisco de Quito) 4 (20).
Cisneros-Heredia, D.F. & Angulo, A. (2009) Cochranella mcdiarmidi. 2009.2 IUCN Red List of Threatened Species.
Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. & Angulo, A. (2009) Centrolene condor. 2009.2 IUCN Red List of Threatened Species.
Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Angulo, A. (2009) Cochranella puyoensis. 2009.2 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. (Ed.) (2009) 2da Cumbre Mundial de Evolución. Aula Magna (Publicación Quincenal de la
Universidad San Francisco de Quito) 4 (18): 5-8.
Cisneros-Heredia, D.F. (2009) Capital Birding: Quito. Neotropical Birding (Birding Magazine of the Neotropical Bird
Club) 4: 40-47.















Cisneros-Heredia, D.F. (2008) Cochranella amelie. 2009.2 IUCN Red List of Threatened Species. Cambridge, United
Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. (2008) Centrolene durrellorum. 2009 IUCN Red List of Threatened Species. Cambridge, United
Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. (2008) Centrolene mariaelenae. 2009 IUCN Red List of Threatened Species. Cambridge,
United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. (2008) Cochranella erminea. 2009 IUCN Red List of Threatened Species. Cambridge, United
Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. (2008) Cochranella resplendens. 2009 IUCN Red List of Threatened Species. Cambridge,
United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. (2008) Hypsiboas nympha. 2009 IUCN Red List of Threatened Species. Cambridge, United
Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. (2008) Nymphargus buenaventura. 2009 IUCN Red List of Threatened Species. Cambridge,
United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. (2008) Nymphargus laurae. 2009 IUCN Red List of Threatened Species. Cambridge, United
Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Jungfer, K.-H., Grant, T., Cisneros-Heredia, D.F., Solís, F., Ibáñez, R., Chaves, G., Savage, J.,
Jaramillo, C., Fuenmayor, Q., Bolaños, F., Lips, K., Lynch, J.D., & Almendáriz, A. (2008) Gastrotheca cornuta. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Heyer, R. & Cisneros-Heredia, D.F. (2008) Leptodactylus peritoaktites. 2009 IUCN Red List of Threatened Species.
Cambridge, United Kingdom. (http://www.iucnredlist.org)
Heyer, R. & Cisneros-Heredia, D.F. (2008) Leptodactylus rhodomerus. 2009 IUCN Red List of Threatened Species.
Cambridge, United Kingdom. (http://www.iucnredlist.org)
Toral, E., Morales, M., Cisneros-Heredia, D.F., Coloma, L.A., Ron, S. & Bustamante, M.R. (2004) Atelopus exiguus.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. & Zak, V. (2000) An Annotated List of the Birds of the Rio Guajalito Protector Forest.
[on line]. Ver. 1.0 (February 20, 2000). Quito, Ecuador.
Cisneros-Heredia, D.F. (1999) The Amphibians and Reptiles of the Tiputini Biodiversity Station. Universidad San
Francisco de Quito: Tiputini Biodiversity Station. Quito.
Cisneros-Heredia, D.F., Ed (1999) An Annotated List of the Birds of the Tiputini Biodiversity Station. Universidad San
Francisco de Quito: Tiputini Biodiversity Station. Quito.
Touzet, J.-M. & Cisneros-Heredia, D.F. (1998) Lianas con Dientes o Serpientes-Palo. Reptinotas Mayo-Junio 1998,
1(3).
Touzet, J.-M. & Cisneros-Heredia, D.F. (1998) Las Serpientes del Género Rhinobothryum. Reptinotas Julio-Agosto
1998, 1(4).
Cisneros-Heredia, D.F. (1997) Inventario Faunistico de la Hacienda La Joya, Prov. de Pichincha, Ecuador.
Fundación Ornitológica del Ecuador, CECIA: Reporte Técnico. Quito.
Cisneros-Heredia, D. F., Ed. (1996) Avifauna No. 4. Fundación Ornitológica del Ecuador, CECIA. Quito.
Cisneros-Heredia, D. F., Ed. (1996) Avifauna No. 3 Año 3. Fundación Ornitológica del Ecuador, CECIA. Quito.
Cisneros-Heredia, D.F. & Cáceres, F. (1995) Lista de aves observadas en los humedales de La Segua.
28/Agosto/1995. Fundación Ornitológica del Ecuador, CECIA: Reporte Técnico. Quito.
Phillips, R. & Cisneros-Heredia, D.F. (1995) Lista Preliminar de Plantas y Animales del Parque Metropolitano de
Quito. Fundación Ornitológica del Ecuador, CECIA: Club de Observadores de Aves. Quito.
Cisneros-Heredia, D.F., Ed. (1995) Avifauna No. 2 Año 2. Fundación Ornitológica del Ecuador, CECIA. Quito.
Cisneros-Heredia, D.F. (1994) Manual: Como Atraer Aves a su Jardín. Fundación Ornitológica del Ecuador,
CECIA. Quito.


















Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Icochea, J., Angulo, A., & Jungfer, K.-H. (2006) Chiasmocleis
anatipes. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. (2005) Trogon personatus, Trogon Enmascarado - Masked Trogon. Boletin SAO (Sociedad
Antioqueña de Ornitología) XV(2): 123-124
Cisneros-Heredia, D.F. (2005) Anfibios y Reptiles, pp. 30-33.
En: Guia EcoRuta El Paseo del Quinde. CORPEI / USFQ. Quito.
Cisneros-Heredia, D.F. (2005) Amphibians and Reptiles, pp. 30-33.
In: Guide EcoRuta El Paseo del Quinde. CORPEI / USFQ. Quito.
Almendáriz A. & Cisneros-Heredia, D.F. (2005) Atelopus sp. 17. En: Rueda-Almonacid, J.V. et al. (Eds.) Ranas
Arlequines. Serie Libretas de Campo 5, Conservacion Internacional. Descarga la guía completa AQUÍ.
Almendáriz A. and Cisneros-Heredia, D.F. (2005) Atelopus sp. 19. En: Rueda-Almonacid, J.V. et al. (Eds.) Ranas
Arlequines. Serie Libretas de Campo 5, Conservacion Internacional. Descarga la guía completa AQUÍ.
Almendáriz, A., Cisneros-Heredia, D.F., & Rueda-Almonacid, J.V. (2005) Atelopus elegans. En: Rueda-Almonacid, J.V.
et al. (Eds.) Ranas Arlequines. Serie Libretas de Campo 5, Conservacion Internacional. Descarga la guía completa
AQUÍ.
Cisneros-Heredia D.F. & Lynch, J.D. (2005) Atelopus guanujo. En: Rueda-Almonacid, J.V. et al. (Eds.) Ranas
Arlequines. Serie Libretas de Campo 5, Conservacion Internacional. Descarga la guía completa AQUÍ.
Cisneros-Heredia D.F. & Rueda-Almonacid, J.V. (2005) Atelopus arthuri. En: Rueda-Almonacid, J.V. et al. (Eds.)
Ranas Arlequines. Serie Libretas de Campo 5, Conservacion Internacional. Descarga la guía completa AQUÍ.
Cisneros-Heredia, D.F. & Lynch, J. D. (2005) Atelopus mindoensis. En: Rueda-Almonacid, J.V. et al. (Eds.) Ranas
Arlequines. Serie Libretas de Campo 5, Conservacion Internacional. Descarga la guía completa AQUÍ.
Cisneros-Heredia, D.F. (2005) Atelopus boulengeri. En: Rueda-Almonacid, J.V. et al. (Eds.) Ranas Arlequines. Serie
Libretas de Campo 5, Conservacion Internacional. Descarga la guía completa AQUÍ.
Galvis P.A. & Cisneros-Heredia, D.F. (2005) Atelopus longirostris. En: Rueda-Almonacid, J.V. et al. (Eds.) Ranas
Arlequines. Serie Libretas de Campo 5, Conservacion Internacional. Descarga la guía completa AQUÍ.


















Almendáriz, A., Cisneros-Heredia, D.F., Coloma, L.A., Ron, S., Wilkinson, W.2004. Caecilia dunni. 2009 IUCN Red
List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Allobates fratisenescus. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Allobates insperatus. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Bolitoglossa equatoriana. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Colostethus fugax. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Osteocephalus alboguttatus.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Osteocephalus deridens. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Osteocephalus fuscifacies. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Pristimantis librarius. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Pristimantis orphnolaimus. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Pristimantis paululus. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Pristimantis trachyblepharis.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H., Coloma, L.A., Ron, S. 2004. Rhinella festae. 2009 IUCN Red
List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Almendáriz, A., Cisneros-Heredia, D.F., Wilkinson, W. 2004. Chthonerpeton onorei. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)




























Angulo, A., Azevedo-Ramos, C., Coloma, L.A., Ron, S., Hoogmoed, M., Gascon, C., Icochea, J. & Cisneros-Heredia,
D.F. (2004) Phyllomedusa palliata. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Angulo, A., Coloma, L.A., Ron, S., Almeida, D., Nogales, F. & Cisneros-Heredia, D.F. (2004) Engystomops
pustulatus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Angulo, A., Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Ceratophrys stolzmanni. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Angulo, A., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Jungfer, K.-H. & Icochea, J. (2004) Hylomantis hulli.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.(http://www.iucnredlist.org)
Angulo, A., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Jungfer, K.-H. & Icochea, J. (2004) Phyllomedusa
coelestis. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Angulo, A., Coloma, L.A., Ron, S., Hoogmoed, M., Castro, F., Rueda, J.V., Cisneros-Heredia, D.F. & Icochea, J.
(2004) Cruziohyla craspedopus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Angulo, A., Coloma, L.A., Ron, S., Jungfer, K.-H., & Cisneros-Heredia, D.F. (2004) Osteocephalus yasuni. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Angulo, A., Coloma, L.A., Ron, S., Jungfer, K.-H. & Cisneros-Heredia, D.F. (2004) Rhinella roqueana. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Angulo, A., Coloma, L.A., Ron, S., Jungfer, K.-H., Cisneros-Heredia, D.F., Castro, F. & Rueda, J.V. (2004)
Osteocephalus verruciger. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Angulo, A., Coloma, L.A., Ron, S., Nogales, F., Cisneros-Heredia, D.F., Neira, D. & Almeida, D. (2004) Leptodactylus
labrosus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Angulo, A., Coloma, L.A., Ron, S., Nogales, F., Cisneros-Heredia, D.F., Almeida, D. & Yánez-Muñoz, M. (2004)
Leptodactylus ventrimaculatus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Azevedo-Ramos, C., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Icochea, J., Angulo, A. & Hoogmoed, M. (2004)
Dendropsophus riveroi. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Azevedo-Ramos, C., Hoogmoed, M., Coloma, L.A., Ron, S., Castro, F., Rueda, J.V., Cisneros-Heredia, D.F., Icochea,
J. & Angulo, A. (2004) Rhinella dapsilis. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)













































Bolívar, W., Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Centrolene scirtetes. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Bolívar, W., Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Hyloscirtus pantostictus. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Bolívar, W., Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Hypsiboas picturatus. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Bolívar, W., Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Rhaebo blombergi. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Bolívar, W., Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Rhaebo hypomelas. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Bolívar, W., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Grant, T. (2004) Osornophryne bufoniformis. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Bolívar, W., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Jungfer, K.-H. (2004) Hyloscirtus alytolylax. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Bolívar, W., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Lynch, J. & Wild, E. (2004) Centrolene balionotum.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Bolívar, W., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Lynch, J. & Jungfer, K.-H. (2004) Hypsiboas pellucens.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Bolívar, W., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Renjifo, J.M. (2004) Gastrotheca guentheri. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)

Bolívar, W., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Wild, E. (2004) Centrolene ballux. 2009 IUCN Red List
of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Bolívar, W., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Wild, E. (2004) Centrolene grandisonae. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Bustamante, M.R., Bolívar, W., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Castro, F., Rueda, J.V., Lötters, S. &
Acosta-Galvis, A. (2004) Atelopus longirostris. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Castro, F. & Cisneros-Heredia, D.F. (2004) Pristimantis parvillus. 2009 IUCN Red List of Threatened Species.
Cambridge, United Kingdom. (http://www.iucnredlist.org)
Castro, F., Herrera, M.I., Azevedo-Ramos, C., Rodríguez, L., Martinez, J.L., Cisneros-Heredia, D.F., Coloma, L.A. &
Ron, S. (2004) Pristimantis martiae. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Castro, F., Herrera, M.I., Coloma, L.A., Ron, S., Lynch, J., Cisneros-Heredia, D.F. & Yánez-Muñoz, M. (2004)
Pristimantis w-nigrum. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Castro, F., Herrera, M.I., Ron, S., Coloma, L.A., Cisneros-Heredia, D.F., Lynch, J. & Yánez-Muñoz, M. (2004)
Pristimantis eremitus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Castro, F., Herrera, M.I., Ron, S., Coloma, L.A., Yánez-Muñoz, M., Lynch, J. & Cisneros-Heredia, D.F. (2004)
Pristimantis quinquagesimus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Castro, F., Ron, S., Coloma, L.A. & Cisneros-Heredia, D.F. (2004) Pristimantis apiculatus. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Castro, F., Ron, S., Coloma, L.A., Cisneros-Heredia, D.F. & Bolívar, W. (2004) Strabomantis anomalus. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)




















Castro, F., Ron, S., Coloma, L.A., Cisneros-Heredia, D.F. & Morales, M. (2004) Pristimantis subsigillatus. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Castro, F., Ron, S., Coloma, L.A., Cisneros-Heredia, D.F., Yánez-Muñoz, M. & Bolívar, W. (2004) Pristimantis
appendiculatus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Castro, F., Ron, S., Coloma, L.A., Morales, M. & Cisneros-Heredia, D.F. (2004) Pristimantis labiosus. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Castro, F., Ron, S., Coloma, L.A., Yánez-Muñoz, M. & Cisneros-Heredia, D.F. (2004) Pristimantis calcarulatus. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F., Almendáriz, A., Coloma, L.A., Ron, S., Wilkinson, M. & Wake, M. (2004) Epicrionops
marmoratus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F., Almendáriz, A., Martinez, J.L. & Neira, D. (2004) Noblella lochites. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F., Almendáriz, A. & Wilkinson, W. (2004) Caecilia abitaguae. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F., Almendáriz, A. & Wilkinson, W. (2004) Caecilia crassisquama. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F., Almendáriz, A. & Wilkinson, W. (2004) Caecilia subterminalis. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F., Almendáriz, A., Yánez-Muñoz, M., Coloma, L.A. & Ron, S. (2004) Ameerega erythromos.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)




















Cisneros-Heredia, D.F., Almendáriz, A., Yánez-Muñoz, M., Coloma, L.A. & Ron, S. (2004) Rhaebo caeruleostictus.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F., Almendáriz, A., Yánez-Muñoz, M., Coloma, L.A. & Ron, S. (2004) Hyloxalus infraguttatus.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F., Almendáriz, A., Yánez-Muñoz, M., Coloma, L.A. & Ron, S. (2004) Hyloxalus toachi. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F. & Jungfer, K.-H. (2004) Dendropsophus carnifex. 2009 IUCN Red List of Threatened Species.
Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F., Morales, M. & Lynch, J. (2004) Pristimantis rosadoi. 2009 IUCN Red List of Threatened
Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F., Yánez-Muñoz, M., Almendáriz, A., Coloma, L.A. & Ron, S. (2004) Bolitoglossa chica. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Cisneros-Heredia, D.F., Yánez-Muñoz, M., Coloma, L.A. & Ron, S. (2004) Atelopus balios. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Almeida, D., Nogales, F., Almendáriz, A., Cisneros-Heredia, D.F., Jungfer, K.-H. & Ron, S. (2004)
Hyloxalus cevallosi. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Coloma, L.A., Lötters, S. & Cisneros-Heredia, D.F. (2004) Atelopus palmatus. 2009 IUCN Red List of Threatened
Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Almeida, D. & Cisneros-Heredia, D.F. (2004) Pristimantis sobetes. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)




















Coloma, L.A., Ron, S., Azevedo-Ramos, C., Bolívar, W., Castro, F., Rueda, J.V. & Cisneros-Heredia, D.F. (2004)
Ecnomiohyla tuberculosa. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Bolívar, W., Cisneros-Heredia, D.F. & Lötters, S. (2004) Atelopus elegans. 2009 IUCN Red
List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Bustamante, M.R., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Atelopus guanujo. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Bustamante, M.R., Lötters, S. & Cisneros-Heredia, D.F. (2004) Atelopus nepiozomus. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Bustamante, M.R., Yánez-Muñoz, M., Cisneros-Heredia, D.F. & Almendáriz, A. (2004)
Atelopus mindoensis. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Centrolene audax. 2009 IUCN Red List of Threatened
Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Centrolene ocellifera. 2009 IUCN Red List of Threatened
Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Centrolene pipilatum. 2009 IUCN Red List of Threatened
Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Hyalinobatrachium pellucidum. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Nymphargus siren. 2009 IUCN Red List of Threatened
Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)




















Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Almeida, D. (2004) Pristimantis orcesi. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Barycholos pulcher. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Dendropsophus gryllatus. 2009 IUCN Red
List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Engystomops coloradorum. 2009 IUCN Red
List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Gastrotheca plumbea. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Osornophryne sumacoensis. 2009 IUCN Red
List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Pristimantis crenunguis. 2009 IUCN Red List
of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Pristimantis ornatissimus. 2009 IUCN Red
List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Pristimantis tenebrionis. 2009 IUCN Red List
of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Strabomantis necerus. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)




















Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Almendáriz, A. & Bolívar, W. (2004) Osornophryne guacamayo. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Almendáriz, A. & Wilkinson, W. (2004) Oscaecilia equatorialis. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Azevedo-Ramos, C., Castro, F., Rueda, J.V., Icochea, J., Heyer, R. &
De la Riva, I. (2004) Leptodactylus discodactylus. 2009 IUCN Red List of Threatened Species. Cambridge, United
Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Castro, F. & Rueda, J.V. (2004) Hemiphractus bubalus. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Grant, T. & Bolívar, W. (2004) Centrolene heloderma. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Icochea, J., Angulo, A. & Jungfer, K.-H. (2004) Bolitoglossa
peruviana. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Icochea, J., Angulo, A., Castro, F., Rueda, J.V. & Bolívar, W. (2004)
Hemiphractus proboscideus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Icochea, J., Angulo, A. & Jungfer, K.-H. (2004) Hyloscirtus
albopunctulatus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Icochea, J., Angulo, A. & Jungfer, K.-H. (2004) Nyctimantis rugiceps.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Jungfer, K.-H. & Bolívar, W. (2004) Scinax quinquefasciatus. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)




















Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Yánez-Muñoz, M. (2004) Andinophryne olallai. 2009 IUCN Red List
of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Yánez-Muñoz, M. (2004) Osornophryne antisana. 2009 IUCN Red
List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Grant, T., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Osornophryne talipes. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Icochea, J., Cisneros-Heredia, D.F., Jungfer, K.-H. & Angulo, A. (2004) Hyloxalus nexipus.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Icochea, J., Jungfer, K.-H. & Cisneros-Heredia, D.F. (2004) Syncope antenori. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Jungfer, K.-H. & Cisneros-Heredia, D.F. (2004) Osteocephalus mutabor. 2009 IUCN Red List
of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Jungfer, K.-H., Cisneros-Heredia, D.F., Almendáriz, A., Solís, F., Ibáñez, R., Jaramillo, C. &
Fuenmayor, Q. (2004) Agalychnis litodryas. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Jungfer, K.-H., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Epipedobates espinosai. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Lynch, J., Cisneros-Heredia, D.F. & Wild, E. (2004) Centrolene peristictum. 2009 IUCN Red
List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Morales, M. & Cisneros-Heredia, D.F. (2004) Pristimantis muricatus. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)




















Coloma, L.A., Ron, S., Morales, M. & Cisneros-Heredia, D.F. (2004) Pristimantis walkeri. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Wild, E. & Cisneros-Heredia, D.F. (2004) Centrolene ballux. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Wild, E., Cisneros-Heredia, D.F., Solís, F., Ibáñez, R., Jaramillo, C., Chaves, G., Savage, J.,
Cruz, G., Wilson, L.W., Köhler, G., Kubicki, B. & Berlin, E. (2004) Cochranella spinosa. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Wild, E., Cisneros-Heredia, D.F., Solís, F., Ibáñez, R., Santos-Barrera, G. & Kubicki, B. (2004)
Hyalinobatrachium fleischmanni. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Yánez-Muñoz, M. & Cisneros-Heredia, D.F. (2004) Andinophryne colomai. 2009 IUCN Red
List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Yánez-Muñoz, M., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Bolitoglossa sima. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Yánez-Muñoz, M., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Epipedobates machalilla.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Yánez-Muñoz, M., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Hyloxalus awa. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Coloma, L.A., Ron, S., Yánez-Muñoz, M., Cisneros-Heredia, D.F. & Almendáriz, A. (2004) Hyloxalus whymperi. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
De la Riva, I., Angulo, A., Azevedo-Ramos, C., Cisneros-Heredia, D.F., Hoogmoed, M., Castro, F. & Rueda, J.V.
(2004) Scarthyla goinorum. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)




















Icochea, J., Coloma, L.A., Ron, S., Almendáriz, A., Cisneros-Heredia, D.F. & Jungfer, K.-H. (2004) Ranitomeya
duellmani. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Icochea, J., Coloma, L.A., Ron, S., Angulo, A., Cisneros-Heredia, D.F., Jungfer, K.-H. & Wilkinson, W. (2004)
Caecilia disossea. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Icochea, J., Coloma, L.A., Ron, S. & Cisneros-Heredia, D.F. (2004) Trachycephalus jordani. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Icochea, J., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Angulo, A. & Jungfer, K.-H. (2004) Gastrotheca longipes.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Icochea, J., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Wilkinson, W. (2004) Caecilia attenuata. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Icochea, J., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Wilkinson, W. (2004) Oscaecilia bassleri. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Icochea, J., Coloma, L.A., Ron, S., Jungfer, K.-H., Angulo, A. & Cisneros-Heredia, D.F. (2004) Allobates zaparo.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Icochea, J., Coloma, L.A., Ron, S., Jungfer, K.-H., Angulo, A. & Cisneros-Heredia, D.F. (2004) Ranitomeya
reticulata. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Icochea, J., Coloma, L.A., Ron, S., Jungfer, K.-H. & Cisneros-Heredia, D.F. (2004) Ameerega parvula. 2009 IUCN
Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Icochea, J., Coloma, L.A., Ron, S., Reichle, S., Angulo, A. & Cisneros-Heredia, D.F. (2004) Gastrotheca testudinea.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Icochea, J., Coloma, L.A., Ron,S., Reichle, S., Castro, F., Rueda,J.V., Angulo,A. & Cisneros-Heredia, D.F. (2004)
Allobates trilineatus.2009 IUCN Red List of Threatened Species.Cambridge,United Kingdom. (http://www.iucnredlist.org)
Kubicki, B., Bolaños, F., Chaves, G., Solís, F., Ibáñez, R., Coloma, L.A., Ron, S., Wild, E., Cisneros-Heredia, D.F. &
Renjifo, J.M. (2004) Centrolene prosoblepon. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Lynch, J., Castro, F., Cisneros-Heredia, D.F. & Wilkinson, W. (2004) Caecilia guntheri. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Lynch, J., Renjifo, J.M., Cisneros-Heredia, D.F., Solís, F., Ibáñez, R., Jaramillo, C., Fuenmayor, Q., Wilkinson, M. &
Wake, M. (2004) Caecilia nigricans. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Rodríguez, L., Martinez, J.L., Azevedo-Ramos, C., Cisneros-Heredia, D.F., Icochea, J. & Hoogmoed, M. (2004)
Pseudopaludicola ceratophryes. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Rodríguez, L., Martinez, J.L., Coloma, L.A., Ron, S., Azevedo-Ramos, C., Castro, F., Rueda, J.V., Cisneros-Heredia,
D.F., Hoogmoed, M. & Gascon, C. (2004) Pristimantis ockendeni. 2009 IUCN Red List of Threatened Species.
Cambridge, United Kingdom. (http://www.iucnredlist.org)
Rodríguez, L., Martinez, J.L., Coloma, L.A., Ron, S., Castro, F., Rueda, J.V., Icochea, J., Angulo, A., Gascon, C. &
Cisneros-Heredia, D.F. (2004) Pristimantis lanthanites. 2009 IUCN Red List of Threatened Species. Cambridge,
United Kingdom. (http://www.iucnredlist.org)
Rodríguez, L., Martinez, J.L., Coloma, L.A., Ron, S., Castro, F., Rueda, J.V., Icochea, J., Angulo, A., Gascon, C. &
Cisneros-Heredia, D.F. (2004) Pristimantis malkini. 2009 IUCN Red List of Threatened Species. Cambridge, United
Kingdom. (http://www.iucnredlist.org)
Rodríguez, L., Martinez, J.L., Coloma, L.A., Ron, S., Castro, F., Rueda, J.V., Cisneros-Heredia, D.F. & Gascon, C.
(2004) Pristimantis acuminatus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Rodríguez, L., Martinez, J.L., Coloma, L.A., Ron, S., Castro, F., Rueda, J.V., Cisneros-Heredia, D.F. & Gascon, C.
(2004) Pristimantis lacrimosus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Rodríguez, L., Martinez, J.L., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F., Angulo, A., Icochea, J. & Acosta, A.
(2004) Hyalinobatrachium munozorum. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom.
(http://www.iucnredlist.org)
Rodríguez, L., Martinez, J.L., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Angulo, A. (2004) Pristimantis
diadematus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Rodríguez, L., Martinez, J.L., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Icochea, J. (2004) Pristimantis
exoristus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Rodríguez, L., Martinez, J.L., Coloma, L.A., Ron, S., Cisneros-Heredia, D.F. & Angulo, A. (2004) Pristimantis
quaquaversus. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Rodríguez, L., Martinez, J.L., Icochea, J., Jungfer, K.-H. & Cisneros-Heredia, D.F. (2004) Pristimantis muscosus.
2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Ron, S., Coloma, L.A., Bustamante, M.R., Cisneros-Heredia, D.F., Almendáriz, A. & Morales, M. (2004) Atelopus
boulengeri. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Ron, S., Coloma, L.A., Bustamante, M.R., Cisneros-Heredia, D.F., Almendáriz, A., Yánez-Muñoz, M. (2004) Atelopus
coynei. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Ron, S., Coloma, L.A., Bustamante, M.R., Cisneros-Heredia, D.F., Lötters, S. & La Marca, E. (2004) Atelopus
planispina. 2009 IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Ron, S., Coloma, L.A., Bustamante, M.R., Cisneros-Heredia, D.F. & Yánez-Muñoz, M. (2004) Atelopus arthuri. 2009
IUCN Red List of Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)
Solís, F., Ibáñez, R., Chaves, G., Savage, J., Bolaños, F., Kubicki, B., Jaramillo, C., Fuenmayor, Q., Coloma, L.A., Ron,
S., Cisneros-Heredia, D.F., Bolívar, W. & Almendáriz, A. (2004) Hyalinobatrachium valerioi. 2009 IUCN Red List of
Threatened Species. Cambridge, United Kingdom. (http://www.iucnredlist.org)











































































